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570<br />

COLLEGE ENTOMOLOGY<br />

cular and nocturnal. The larvre of a number of species including Calophus angustus<br />

Lee. and Nacerda melanura (Linn.) breed in dead coniferous timbers.<br />

The latter has become quite injurious to lower. shaded, and damp timbers of<br />

wharves along the California coast. Copidita 4-maculata (Mots.) breeds in old<br />

wet mine and bridge timbers in California.<br />

Family MORDELLIDE (Leach 1815).<br />

Tumbling Flower Beetles. A large family of small beetles; characterized by<br />

laterally compressed, arched, silky bodies pointed posteriorly and extending beyond<br />

the tips of the elytra; long legs; and jumping and tumbling reactions when<br />

disturbed. The adults are common on flowers. There are about 800 species<br />

distributed throughout the world.<br />

Family MELOIDlE Thomson 1859 (Me-lo'i.-dre, from the new Latin MeloiJ;<br />

origin uncertain). German. Pflasterldifer, 6lkafe.r, Maiwiirmer. French,<br />

Meloides. Blister Beetles, Oil Beetles, Meloids.<br />

Medium-sized, elongated, somewhat cylindrical or robust, soft-bodied beetles<br />

which are characterized in some species by having short, loosely connected<br />

elytra. Smooth, rugose, sculptured, punctate, pubescent. or hairy. Colors<br />

mostly somber - black, gray, brownish. or tan - or bright, with metallic<br />

iridescences. Some species are beautifully sculptured, brilliantly colored. and<br />

grotesquely formed. Head hypognathous, large, free, constricted at base.<br />

Eyes large, widely separated. Antennre sedform or certain segments enlarged<br />

in males; ll-segmented. Mouth parts well developed. Prothorax small, usually<br />

not much wider than the head, much narrower than rest of thorax, and scarcely<br />

longer than wide. Wings well developed, abortive, or absent. Elytra longer or<br />

shorter than the body. not closely joined in the middle, even overlapping, and<br />

widely divergent apically. Legs long and slender. Tarsi long, five-segmented.<br />

Anterior and middle cOXa! large and contiguous. Abdomen often with the posterior<br />

segments exposed and with six visible siemites.<br />

These beetles have a complex development known as hypermetamorphosis.<br />

The very minute eggs are laid in great numbers in masses in the soil. The newly<br />

hatched young, called primary larvre or triungulins, are minute and campodeiform<br />

with well-developed legs, antennre, and cerci. They are very active and<br />

seek out egg masses or brood chambers of ground-nesting insects, or they may<br />

actually attach themselves to certain adult hosts and ride to the nests or brood<br />

chambers of the latter. Here they feed upon the honey or food stored" for the<br />

host larvre or are predacious upon the latter. But very shortly they transform<br />

into apodous, sluggish, scarabreoid type of larvre and remain until fully developed,<br />

passing, in some cases, into still a third type of larval and into a I;lrepupa,<br />

a pupa, and finally into an adult.<br />

The charming sketch by Fabre (1857) describes the life history of the European<br />

Sitart's muralis Forster, a small black and yellow beetle 8-10 mm. long<br />

which develops in nests of bees belonging to the genus Anthophora. The eggs<br />

are laid near the ground nests of the bees during late summer and hatch in the

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