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520<br />

COLLEGE ENTOMOLOGY<br />

The larva:! of many species pupate in a cell in the ground, burrow in wood, or<br />

, other secluded places; while certain leaf weevils spin lace-like cocoons from a<br />

sticky quick-drying fluid taken from the anal opening by the mandibles and<br />

slowlY and skillfully woven. The normal life cycles may be very short, requiring<br />

from one to four generations a year, or may be quite long, involving from<br />

one to five years for a single complete life history. Rarely from 25 to 30 years<br />

have been recorded for certain wood-boring beetles. Hibernation and restivation<br />

may occur in all the various stages.<br />

The fossil record may be summarized as follows:<br />

Order PROTOCOLEOPTERA Tillyard 1924, from Upper Permian, Belmont,<br />

New South Wales. Early ancestors. .<br />

Trias, Upper - Ipswich, Australia - Beetles abundant.<br />

Liassic - Europe - abundant.<br />

Jurassic, Upper - Europe - abundant.<br />

Tertiary - North America - abundant.<br />

Eocene - many modern genera occur.<br />

Baltic Amber - nearly 500 species known.<br />

Miocene - Florissant, Colo. - nearly 1,000 species known.<br />

BASIS FOR CLASSIFICATION FOR COLEOPTERA<br />

1. Character of the Exoskeleton and Elytra: Whether hard and heavily scleratized<br />

or soft and leather-like; the surface, whether smoothed and polished<br />

or pubescent, hairy, scaly, punctured, pitted, rugose, ribbed, or striated;<br />

length of elytra, whether covering or only partially covering the abdomen.<br />

The wing venation is also used to some extent in separating families.<br />

2. Antennre: The shape of the antenna:! is exceedingly variable and may be<br />

setiform or awl-like and filiform or thread-like, in which all segments are<br />

more or less alike and gradually diminish in size to a point apically;<br />

moniliform, or with the segments rounded and bead-like; serrate, or with<br />

the segments extending laterally on the outside like saw-teeth; clavate, in<br />

which the segments gradually increase in size apically to form a club;<br />

capitate, ending in usually three to five large loose or fused segments which<br />

form a terminal enlargement or ball-like structure; geniculate or elbowed,<br />

consisting of a small basal segment and the longest segment forming the<br />

!'lcape, followed by a short pedicel and a number of small ring-joints and<br />

segments composing the flagellum (these segments are often held at an<br />

angle to the scape); lamellate geniculate or capitate types, in which the<br />

apical portion is composed of a number of lamellre that fit together like<br />

leaves of a book; pectinate, where the segments have long lateral slender<br />

extensions so as to produce a comb-like appearance and occurring usually<br />

in males. The position of the insertion of the antennre on the head is also<br />

important.<br />

:1. Abdominal Stemites: In these closely joined insects there is a fusion of<br />

segments at the base and apex of the abdomen with the result that the

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