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THE ANATOMY OF INSECTS 39<br />

Organs of sound and hearing. - It is generally supposed that insects capable of<br />

producing sound are also able to hear. Organs of sound have previously been<br />

discussed in the chapter on external anatomy and consist of many kinds of<br />

stridulating organs, flight mechanisms, tympanums or drums, and other devices.<br />

The chordotonal organs are variable complex structures which are sound<br />

perceptors or tension receptors and according to Snodgrass (1935) are cellular<br />

elements with or without scolops or sense rods which constitute elongated structures<br />

both of whose ends are attached to the cuticle. johnston's organ, first observed<br />

by Christopher Johnston in 1855, is a specialized cup-like chordotonal<br />

organ occupying much of the second antennal segment in DIPTERA, being<br />

especially well developed in hairy antennre of male mosquitoes and midges<br />

and, to a lesser degree, of bluebottle and greenbottle flies; in LEPIDOPTERA<br />

in satyrid butterflies, and in HYMENOPTERA in paper wasps (Imms). The<br />

tympanal auditory organs on each side of abdominal segment I of grasshoppers,<br />

and on the fore tibire of crickets and katydids, are highly specialized structures<br />

known as the "ears" of these insects.<br />

GLANDULAR SYSTEM<br />

The many variable and important glandular organs of insects are not closely<br />

enough coordinated to constitute a "system" in the real sense of the word, so<br />

this term is used in a very general sense to denote a grouping of miscellaneous<br />

glandular organs of similar structures and functions.<br />

Salivary glands - a variable number located in the head and thoracic regions<br />

which open by means of one to four or more pairs of ducts at the base of the<br />

hypopharynx of adult and larval forms.<br />

The frontal gland of the nasute termite exudes a sticky fluid used as a defensive<br />

and offensive weapon against enemies.<br />

Molting glands - numerous minute unicellular bodies, secreting fluids necessary<br />

for the shedding of the skin during the process of molting in insects exposed<br />

to air, light, and heat.<br />

Silk glands - often a pair of very large tubes, occupying much of the body<br />

cavities of lepidopterous larvre. The silk is drawn through a single spinneret on<br />

the labium. The glands of Filippi are associated with the silk prlfss at the posterior<br />

region of the mouth. In embiids, silk glands and a series of spinnerets occur<br />

in the enlarged first tarsal segment. In certain curculionid and neuropteroid<br />

larvre silk is produced by the Malpighian tubules, obtained from the anus by<br />

means of the mouth parts, and used to construct lace-like cocoons or for other<br />

purposes. The glands of Batelli are hypodermal glands opening into the rectum<br />

and secreting a fluid which is forced from the anus along with air bubbles to<br />

form the froth or spittle of spittle bugs.<br />

Setal glands are very numerous in the larVa! and adults of many insects.<br />

They are small glands with outlets through lumina or tubes of set:e or "tenant<br />

hairs." The venomous stinging and poisonous hairs of certain "woolly" caterpillars<br />

are of this type. Tenant hairs that secrete sticky fluids also occur on the

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