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HEMIPTERA<br />

thrips, aphids, and leafhoppers, on eggs and newly hatched caterpillars of the<br />

corn ear worm and other moths, and on other small animals. and are at times<br />

of some importance in reducing the numbers of these pests. O. sauten' (Pop.<br />

pius) feeds upon the mulberry leafhopper in Japan.<br />

Family MIRIDlE 1 Hahn 1831 (Mir'i-dre. from the Latin mirus. wonderful).<br />

German, Weichwanzen. Plant Bugs, Leaf Bugs, Capsids, Mirids.<br />

Small to medium-sized, ovate, oblong, or slender, somewhat flattened,<br />

fragile, active species with soft integument, smooth, shiny. pubescent or hairy<br />

bodies, and black, dull, or brilliant colors. Head small, often sharply produced.<br />

Antennre well developed, four-segmented, slender. Eyes well developed.<br />

Ocelli absent. Rostrum long, four-segmented; first segment as long as or<br />

longer than head. Prothorax with or without an apical stricture. Hemelytra<br />

opaque or hyaline, with clavus, corium, cuneus, membrane, and linear un·<br />

differentiated embolium; membrane with two basal cells or areoles and one<br />

longitudinal anal vein; cuneus a well-developed triangular area between the<br />

outer membrane and inner portion. Macropterous, brachypterous, and apterous<br />

forms occur within the genus or even the species. Males are usually macropterous<br />

in species with brachypterous and apterous females. Legs long and<br />

slender. Tarsi normally three-segmented, rarely two-segmented. Claws thick<br />

or slender, widely or sharply curved; with or without bristle-like arolia and<br />

pseudarolia which are of value in separating the subfamilies. Female genitalia<br />

asymmetrical; with sickle-like ovipositor. Male genitalia much used in taxonomic<br />

studies.<br />

The members are largely herbivorous, but many species are distinctly<br />

predacious and a few are both plant and animal feeders. Many of the plantinfesting<br />

forms are most injurious to native vegetation and to cultivated<br />

crops and may fairly swarm over the hosts. The eggs are generally inserted<br />

into the hard and soft tissues of living plants. The adults may hibernate and<br />

emerge in great numbers in the spring. A considerable number of species.<br />

including the tarnished plant bug, Lygus pratensis (Linn.), and Creontiades<br />

pallidus Rambur. carry and transmit plant diseases which may result in large<br />

losses to crops. Predacious species like Cyrtarhinus mundulus Bred. which<br />

feeds on the eggs of the sugar-cane leafhopper in the tropical Pacific area,<br />

and the well-known aphid destroyer, Derreocoris ruber (Linn.), of Europe,<br />

destroy noxious insects and are considered beneficial.<br />

The family is the largest in the suborder. It is usually divided into nine to<br />

16 subfamilies and comprises over 600 genera and 5,000 species. It reaches its<br />

highest development in the Holarctic region. where about 1,100 species are<br />

known. Some of the most important genera are Adelphocl)7is Reuter, Atrac.to·<br />

I The members of this family were originally grouped with the COREID,£ by Leach 1815,<br />

Samouelle 1819. and Stephens 1829. Other designations were CAPSINI Burmeister 1835:<br />

CAPSOIDES Spinola 1837; CAPSIDJE Kirby 1837, Westwood 1840, Butler 1923, Schrllder<br />

1925, Weber 1930, lroms 1934, and others; MIRIDtE Hahn 18.31, Kirk.aldy 1894. Reuter<br />

1910. Van Duzee 1917, Knight 1923. Brues and Melander 1932. Hedicke 1935; PHYTO­<br />

COREIDJE Fieber 1851.

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