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CHAPTER XVIII<br />

15. Order EMBIOPTERAI Shipley 1904<br />

(Em'bi-op'ter-a, from the Greek €fJ,f3ws, lively, + 7r'Tfpa,<br />

wings.) German, Embien. French, Embiides.<br />

Embiids, Embiopterans, Webspinners.<br />

Small, '>lender, campodeiform insects with biting and chewing mouth parts and simple<br />

or hemimetabolous metamorphosis. Head large, antennre filiform; compound eyes<br />

small in female, often large in male; ocelli absent; thorax nearly as long as abdomen;<br />

legs short and stout for running, tarsi three-segmented, first tarsal segment of forelegs<br />

enlarged to contain glands and spinnerets. Wings, two pairs, membranous, similar in<br />

size, shape, and venation, held fiat over body in repose, present or absent in males,<br />

absent in females.<br />

This is a small group of truly peculiar insects which is widely distributed<br />

throughout the tropical and more temperate regions of the world. Because of<br />

their secluded habits, embiids are scarcely known even to entomologists.<br />

Wherever the insects occur, their presence is indicated by the numerous filmy<br />

white webbed tunnels and coverways which they spin copiously throughout the<br />

range of their activities. Sometimes the webs are evident in the debris on the<br />

surface of the soil at the bases of grasses, weeds, and other plants during the<br />

rainy season, but more often they appear to occur under stones, bark, pieces of<br />

wood, dried dung, and similar objects which serve as covering and protection<br />

for their habitations. Some species, like Clothoda uriclti (Sauss.) of Trinidad,<br />

live under the bark of trees and others are reported to inhabit by chance the<br />

nests of ants and termites. In California, where there is a distinct wet winter<br />

season and a dry summer season, colonies of Gynembia tarsalia Ross may be<br />

observed on open, grassy slopes and hillsides during the winter after the first<br />

soaking rains and until the vegetation dries up in early summer, when the<br />

species retreats into the soil. The members of this species are decidedly gregarious<br />

and may occur in large societies; and their silken coverways and tunnels<br />

extend over much of the surface of the ground and of whatever objects they are<br />

living beneath. Although much study and attention has been given to them<br />

over a period of many years, and large numbers have been confined in the<br />

I The order name of these insects has been repeatedly changed. Among the important<br />

names used are: EMBIDINA Hagen 1861. EMBIDOPTERES Lameere 1900. EMBIIDINA<br />

Enderlcin 1903, EMDIODEA Kusnezow 1903, EMBIOIDEA, EMBIARIA IIandlirsch 1903,<br />

EMBIOPTERA Shipley 1904, ADENOPODA Verhoff 1904, OLIGONEURA Bomer 1904,<br />

EMBIJE Jacobson and Bianchi 1905, EMBIDOS Navas 190[;. EMBIDARIA HandUrsch<br />

1906, and MTIOPTERA Enderlein 1912. While the name EMBIIDINA appears to have<br />

priority, the author is using the generally accepted term EMBIOPTERA as recently employed<br />

by Imms 1930, Metcalf and Flint 1932, Folsom and Wardle 1934, DaVIS 1939, and<br />

Ross 1940.<br />

177

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