A synopsis of the South Asian fishes referred to Puntius - Verlag Dr ...

74
100
100
74
100
71
10 changes
100
100
91
92
Garra ceylonensis (LK)
Garra mullya (IN)
Labeo dussumieri (LK)
Labeo dussumieri (IN)
100
83
91
100
Osteochilichthys nashii (IN)
Gonoproktopterus jerdoni (IN)
Gonoproktopterus curmuca (IN)
Tor khudree (LK)
Barbonymus schwarnenfeldii (AQ)
92
92
92
99
50
100
87
99
59
100
100
100
57
100
92
96
87
100 Pethia bandula (LK)
99
98 Pethia nigrofasciata (LK)
100
98
93
100
85
Pethia reval (LK)
Pethia cumingii (LK)
Pethia melanomaculata (LK)
Pethia conchonius (AQ)
Pethia ticto (IN)
Dravidia fasciata (IN)
Dawkinsia assimilis (IN)
Dawkinsia filamentosa (IN)
Dawkinsia singhala (LK)
Dawkinsia srilankensis (LK)
60
99
50
64
64
100
84
100
96
100
64
100
100
100
100
Puntius mahecola (IN)
100
66
Puntius bimaculatus (LK)
Puntius cf. bimaculatus (LK)
Puntius cf. titteya (LK)
Puntius titteya (LK)
Puntius thermalis (LK)
Puntius chola (IN)
Puntius sophore (IN)
Puntius dorsalis (LK)
100
65
Puntius dorsalis (IN)
Puntius kelumi (LK)
Puntius layardi (LK)
Systomus martenstyni (LK)
Systomus sp. ‘Richmondi’ (LK)
Systomus sarana (IN)
100
73
100
100
Systomus timbiri (LK)
Systomus spilurus (LK)
Systomus pleurotaenia (LK)
Outgroup
Last unbranched dorsal-fin
ray serrated; infraorbital 3
deep; SL < 80 mm; rostral
barbels absent; post-epiphysial
fontanelle absent.
Last unbranched dorsal-fin
ray smooth; SL < 80 mm;
rostral, maxillary barbels
present; post-epiphysial
fontanelle absent.
Last unbranched dorsal-fin
ray smooth; rostral barbels
absent; 3 black bars on flank
in juveniles; post-epiphysial
fontanelle absent.
Last unbranched dorsal-fin
ray smooth; 7 branched
dorsal-fin rays; rostral
barbels absent; postepiphysial
fontanelle absent.
Last unbranched dorsal-fin
ray smooth; 8 branched
dorsal-fin rays; rostral barbels
absent; post-epiphysial
fontanelle present.
Last unbranched dorsal-fin
ray serrated; rostral, maxillary
barbels present; SL >
80 mm; 27−34 lateral-line
scales; post-epiphysial
fontanelle absent.
Fig. 1. Bayesian phylogram of the 31 putative species of South Asian Puntius s. l. analysed, with key shared
morphological and osteological characters for each group. See Table 1 for list of material. Provenance: AQ,
aquarium trade; IN, India; LK, Sri Lanka. Posterior probabilities are shown above nodes, parsimony bootstrap
values below nodes. Photographs illustrate species within the respective genera: from top, Pethia bandula, Dravidia
fasciata, Dawkinsia srilankensis, Puntius titteya, Puntius dorsalis and Systomus asoka.
Pethiyagoda et al.: Synopsis of South Asian Puntius