Nicola Arndt und Matthias Pohl - Neobiota
Nicola Arndt und Matthias Pohl - Neobiota Nicola Arndt und Matthias Pohl - Neobiota
secondary in origin but include stands on ancient woodland sites and old plantations dating from the 18 th and 19 th centuries. During the early part of the 20 th century a policy of afforestation was introduced and this gained momentum during the last 4 decades of the century when extensive areas of marginal land were planted. The majority of these plantations consist of introduced coniferous species, principally sitka spruce (Picea sitchensis) 1 and lodgepole pine (Pinus contorta) which grow rapidly on the mostly poor land – principally upland soils, blanket peats and heavy gleys – selected for afforestation. Today, forest and woodland covers 626,000 ha or c. 9.0 % of the land area and the national policy is to increase the forest estate to 17 % of the land area by 2030 (ANON. 2000a). 2 The character of existing native Irish woodlands O’SULLIVAN (1998) estimated that not more than 100,000 ha, or 1.5 % of the land area, consists of broadleaf woodland – including scrub, mostly in stands of less than 100 ha and many very much smaller. More recent surveys indicate that there are c. 77,000ha of native woodland, excluding scrub (HIGGINS et al 2004). Most of these stands are of native species, although they include beech (Fagus sylvatica) and other non-native species. Despite their size they are very important as a reservoir of native species and are recognised as a major repository of biodiversity. They can be broadly categorised as follow (CROSS 1987a): • Remnants of the original forest (ancient woodland), mostly confined to the poorest sites, greatly modified and often over-exploited or largely abandoned silviculturally. • Old plantations, 150 to 200 years old, often with beech, larch (Larix decidua), Scots pine (Pinus sylvestris) and other exotic species, and probably with foreign provenances of native species. • Secondary woodland of relatively recent origin on abandoned farmland and cutaway bog, often scrub-like and with the better quality timber commonly selectively removed. • Recent plantations, mostly of ash, oak and beech. The species complement of native Irish woodlands is considerably poorer than other European countries because of the separation of the island from the continental land mass relatively early after the last glaciation (Table 1). Oak (Quercus species), ash (Fraxinus excelsior), birch (Betula species) and alder (Alnus glutinosa) are the commonest canopy-forming species and hazel (Corylus avellana), holly (Ilex aquifolium), hawthorn (Crataegus monogyna) and sally (Salix cinerea ssp. oleifolia) are the commonest understorey species. Several species which occur in Britain and the nearby continent are absent, most notable of which are important components of the forest climaxes, such as beech, hornbeam (Carpinus betula), lime (Tilia species), sycamore (Acer pseudoplatanus) and field maple (A. campestre). Some of these have been widely planted and regenerate spontaneously, especially beech and sycamore. Scots pine, formerly widespread, is generally considered to be extinct as a native species, although it may have survived until c. 1,000 years ago (BUDD & VON ENGELBRECHTEN 1999). It is now widespread as a result of its re-introduction 200-300 years ago, mostly from Scotland, and it regenerates freely on suitable soils. Certain shrub species, common in forests elsewhere in western Europe, are very local or rare within Irish forests, e.g. Frangula alnus, Prunus padus. The herbaceous flora is also impoverished, with species such as Mercurialis perennis very rare and possibly not native. In contrast ash, which in drier climates thrives principally only in valleys and other moist sites, is 1 Nomenclature for plants follows TUTIN et al. (1964-1980); for animals EU Council Directive 92/43/EEC final version. 362
widespread and an important component of woodlands on base-rich soils. Numerous Atlantic species, such as Hyacinthoides non-scripta, Dryopteris aemula and Salix cinerea ssp. oleifolia, are also widespread and abundant. Table 1: Tree and shrub species considered native to Ireland (after SCANNELL & SYNNOTT 1987). Major components of the canopy and/or shrub layer Alnus glutinosa Betula pubescens B. pendula Corylus avellana Crataegus monogyna Fraxinus excelsior Ilex aquifolium Prunus spinosa Quercus robur Q. petraea Sambucus nigra Salix cinerea ssp. oleifolia Sorbus aucuparia Minor components of the canopy and/or shrub layer Euonymus europaea Malus sylvestris Populus tremula Prunus avium Salix aurita 3 Potential Natural Forest types of Ireland S. caprea Taxus baccata Ulmus glabra Viburnum opulus Rare or local species Arbutus unedo Cornus sanguinea Franguls alnus Juniperus communis Ligustrum vulgare Prunus padus Rhamnus cathartica Salix cinerea ssp. cinerea S. myrsinifolia S. pentandra S. phylicifolia S. purpurea Sorbus anglica S. aria agg. S. rupicola In many countries the maps of potential natural vegetation are the product of an amalgamation of units mapped at a large scale. In contrast, in Ireland no detailed systematic survey or mapping of the vegetation at a large scale has been undertaken. The map of potential natural vegetation for Ireland, as it appears in the 1 : 2,500,000 Map of the Natural Vegetation of Europe, is one of the first attempts to map Irish vegetation in this detail and in this respect it is still a coarse instrument (CROSS 1998). Nonetheless, the map has already stimulated considerable interest and with the associated edaphic and climatic data and the information on the natural vegetation occurring in complex, it provides a useful tool for applications in forestry, nature conservation and planning. Nine forest units are mapped and described, while the degraded raised bogs may be considered a tenth unit. In addition there are several azonal types, covering small areas, which cannot be mapped at this scale (CROSS 1987b, KELLY & IREMONGER 1997). These units, with abbreviated titles, are listed in Table 2; the numbering follows that used in the legend to the map (BOHN et al. 2003). 4 Potential natural vegetation and protected areas There are c. 6,309 ha of protected native woodlands in the country. Of this approximately 1,730 ha occur in 5 national parks, c. 2,359 ha in 32 nature reserves and an additional 2,220 ha in other Natural Heritage Areas (NHAs – sites proposed for protection under national legislation) and proposed Special 363
- Seite 314 und 315: 7 Die Quantifizierung der Ökosyste
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- Seite 321 und 322: eine Verbreitungskarte erstellt wer
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- Seite 337 und 338: Anwendung und Auswertung der Karte
- Seite 339 und 340: Abb. 1: Klimadiagramme ausgewählte
- Seite 341 und 342: (S. graeca, S. hajastana u. a.) und
- Seite 343 und 344: Die Kräuterfluren besiedeln vorwie
- Seite 345 und 346: Picea orientalis gehört zur Sektio
- Seite 347: Anschrift des Autors: Prof. Dr. Gio
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- Seite 370 und 371: Table 3: Principal potential forest
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- Seite 377 und 378: Figure 3: Woodlands and their repla
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- Seite 389 und 390: Im vorliegenden Beitrag steht die V
- Seite 391 und 392: Abb. 1: Birken-Pionierwald mit eind
- Seite 393 und 394: Wollgras- Torfmoos-KI-Typ naß Torf
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- Seite 399 und 400: BOHN, U.; GOLLUB, G. & HETTWER, C.
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secondary in origin but include stands on ancient woodland sites and old plantations dating from the<br />
18 th and 19 th centuries. During the early part of the 20 th century a policy of afforestation was<br />
introduced and this gained momentum during the last 4 decades of the century when extensive areas of<br />
marginal land were planted. The majority of these plantations consist of introduced coniferous species,<br />
principally sitka spruce (Picea sitchensis) 1 and lodgepole pine (Pinus contorta) which grow rapidly on<br />
the mostly poor land – principally upland soils, blanket peats and heavy gleys – selected for<br />
afforestation. Today, forest and woodland covers 626,000 ha or c. 9.0 % of the land area and the<br />
national policy is to increase the forest estate to 17 % of the land area by 2030 (ANON. 2000a).<br />
2 The character of existing native Irish woodlands<br />
O’SULLIVAN (1998) estimated that not more than 100,000 ha, or 1.5 % of the land area, consists of<br />
broadleaf woodland – including scrub, mostly in stands of less than 100 ha and many very much<br />
smaller. More recent surveys indicate that there are c. 77,000ha of native woodland, excluding scrub<br />
(HIGGINS et al 2004). Most of these stands are of native species, although they include beech (Fagus<br />
sylvatica) and other non-native species. Despite their size they are very important as a reservoir of<br />
native species and are recognised as a major repository of biodiversity. They can be broadly<br />
categorised as follow (CROSS 1987a):<br />
• Remnants of the original forest (ancient woodland), mostly confined to the poorest sites, greatly<br />
modified and often over-exploited or largely abandoned silviculturally.<br />
• Old plantations, 150 to 200 years old, often with beech, larch (Larix decidua), Scots pine (Pinus<br />
sylvestris) and other exotic species, and probably with foreign provenances of native species.<br />
• Secondary woodland of relatively recent origin on abandoned farmland and cutaway bog, often<br />
scrub-like and with the better quality timber commonly selectively removed.<br />
• Recent plantations, mostly of ash, oak and beech.<br />
The species complement of native Irish woodlands is considerably poorer than other European<br />
countries because of the separation of the island from the continental land mass relatively early after<br />
the last glaciation (Table 1). Oak (Quercus species), ash (Fraxinus excelsior), birch (Betula species)<br />
and alder (Alnus glutinosa) are the commonest canopy-forming species and hazel (Corylus avellana),<br />
holly (Ilex aquifolium), hawthorn (Crataegus monogyna) and sally (Salix cinerea ssp. oleifolia) are the<br />
commonest <strong>und</strong>erstorey species. Several species which occur in Britain and the nearby continent are<br />
absent, most notable of which are important components of the forest climaxes, such as beech,<br />
hornbeam (Carpinus betula), lime (Tilia species), sycamore (Acer pseudoplatanus) and field maple<br />
(A. campestre). Some of these have been widely planted and regenerate spontaneously, especially<br />
beech and sycamore. Scots pine, formerly widespread, is generally considered to be extinct as a native<br />
species, although it may have survived until c. 1,000 years ago (BUDD & VON ENGELBRECHTEN 1999).<br />
It is now widespread as a result of its re-introduction 200-300 years ago, mostly from Scotland, and it<br />
regenerates freely on suitable soils. Certain shrub species, common in forests elsewhere in western<br />
Europe, are very local or rare within Irish forests, e.g. Frangula alnus, Prunus padus. The herbaceous<br />
flora is also impoverished, with species such as Mercurialis perennis very rare and possibly not native.<br />
In contrast ash, which in drier climates thrives principally only in valleys and other moist sites, is<br />
1 Nomenclature for plants follows TUTIN et al. (1964-1980); for animals EU Council Directive 92/43/EEC final<br />
version.<br />
362