Nicola Arndt und Matthias Pohl - Neobiota

Nicola Arndt und Matthias Pohl - Neobiota Nicola Arndt und Matthias Pohl - Neobiota

22.02.2013 Aufrufe

The deserts of Artemisia lerchiana are widely distributed in various environments on plains with loamy, sandy-loamy and sandy soils (Table. 1). Large areas in this region are covered by communities of sand wormwood (A. arenaria) confined to loose sands. In these communities participation of shrubs is appreciable. Communities of black wormwood (A. pauciflora) are spread on solonetz and saline soils. Often these communities consist only of one species or include some annuals. On solonchak- solonetz they are codominated by Camphorosma monspeliaca. On the right bank of the Ural River (and eastwards) large areas are covered by perennial saltwort deserts on saline soils; these are monodominant communities of Anabasis salsa and communities of Atriplex cana. Table 1: Ecological-phytocoenotical dominant-determinant classification of European deserts. Subzone on the European Vegetation Map Northern lowland plain deserts Southern foothill deserts Formation Ecotype Association, complex of communities Artemisia lerchiana pl 1 Caspian Lowland Artemisia lerchiana + pl hal Artemisia lerchiana-nabasis aphylla + hps Artemisia lerchiana-Poa bulbosa + hps Artemisia lerchiana-Stipa sareptana + hps Artemisia lerchiana-Stipa lessingiana + ps Artemisia lerchiana with Calligonum aphyllum + ps hal Artemisia lerchiana with Calligonum aphyllum and Krascheninnikovia ceratoides + Artemisia pauciflora pl hal Artemisia pauciflora + pl hal Artemisia pauciflora- Camphorosma monspeliaca + Artemisia arenaria ps Artemisia arenaria-Calligonum aphyllum + Anabasis salsa pl hal Anabasis salsa + Atriplix cana pl hal Atriplex cana + Artemisia fragrans pl Artemisia fragrans-Poa bulbosa, Catabrosella humilis Artemisia szowitsiana pl hal Artemisia szowitsiana-annual saltworts Salsola nodulosa pl hal Salsola nodulosa-Poa bulbosa, Catabrosella humilis with Artemisia fragrans Salsola ericoides pl hal Salsola ericoides-Poa bulbosa, Catabrosella humilis with Artemisia fragrans Salsola dendroides pl hal Salsola dendroides-Poa bulbosa, Catabrosella humilis with Artemisia fragrans Eastern Transcaucasus 1 pl − pelitophytic, pl hal − pelitophytic halophytic, hps − hemipsammophytic, hps hal − hemipsammophytic halophytic, ps − psammophytic, ps hal − psammophytic halophytic Solonchak depressions are rather widely distributed in the Caspian Lowland. Around sors (solonchaks lacking vegetation) vegetation cover is formed by communities of hyperhalophytic annual saltworts and perennial saltworts, belonging mostly to the following formations: Halocnemum strobilaceum, Kalidium foliatum, Atriplex verrucifera, A. cana (Table 2). 154 + + + + +

The deserts of the eastern Transcaucasus Region differ from those of the northern Caspian Region by a set of formations (see Table 1). They include the communities of two wormwood species (Artemisia fragrans and A. szowitsiana) and of three perennial saltworts (Salsola nodulosa, S. ericoides and S. dendroides). Ephemeroid grasses (Poa bulbosa and Catabrosella humilis) constantly occur in these communities. On solonchaks in the eastern Transcaucasus, communities of several formations (Halostachys belangeriana, Suaeda microphylla, Gamanthus pilosus, Zeidlitzia florida) are characteristic in addition to the communities of the same formations as in the northern Caspian Region (Halocnemum strobilaceum, Atriplex verrucifera and annual saltworts) (Table 2). The vegetation of the northern Caspian Region is species-poor and represented by a small number of ecotypes correlated with the low diversity of environmental conditions in the area. The Vegetation Map of Europe shows exactly the peculiarities of their distribution. To the west of the Volga River, extensive plains with sandy-loamy soils are occupied by hemipsammophytic deserts of the Artemisia lerchiana-Poa bulbosa type which is locally combined with psammophytic types on sand: Artemisia lerchiana-Calligonum aphyllum and Artemisia arenaria- Calligonum aphyllum with Krascheninnikovia ceratoides. The pelitophytic desert complex is distributed along the steppe boundary as a narrow strip. Depending on salt concentration, the communities of Artemisia lerchiana (on soils with lower salt concentrations) and those of Artemisia pauciflora (on soils with higher salt concentrations) predominate alternately in complexes. Table 2: Formations on solonchak. Formation Northern deserts in the Caspian Lowland Atriplex cana + Kalidium foliatum + Southern foothill deserts in eastern Trans-caucasus Halocnemum strobilaceum + + Atriplex verrucifera + + Annual saltworts + + Kalidium сaspicum + Halostachys belangeriana + Suaeda microphylla + Gamanthus pilosus + Zeidlitzia florida + On the Volga-Ural interfluve psammophytic deserts predominate. In terms of the area covered, the dominant communities are those of Artemisia lerchiana with grasses (Stipa sareptana, S. lessingiana) or psammophytic shrubs and semishrubs (Calligonum aphyllum, Krascheninnikovia ceratoides) on flat sands, stabilized dunes and plains with sandy soils. Communities of Artemisia arenaria are widely distributed, with constant participation of Calligonum aphyllum restricted to semi-moving sands on hills and ridges. Locally, shrub deserts (Calligonum aphyllum, Tamarix ramosissima, T. laxa) are found on the weakly stabilized sands of barkhans. The hemipsammophytic Artemisia lerchiana-Poa bulbosa communities are characteristic of areas on both sides of the Volga River, but their ranges are smaller than those of psammophytic types. Complex 155

The deserts of Artemisia lerchiana are widely distributed in various environments on plains with<br />

loamy, sandy-loamy and sandy soils (Table. 1). Large areas in this region are covered by communities<br />

of sand wormwood (A. arenaria) confined to loose sands. In these communities participation of shrubs<br />

is appreciable. Communities of black wormwood (A. pauciflora) are spread on solonetz and saline<br />

soils. Often these communities consist only of one species or include some annuals. On solonchak-<br />

solonetz they are codominated by Camphorosma monspeliaca.<br />

On the right bank of the Ural River (and eastwards) large areas are covered by perennial saltwort<br />

deserts on saline soils; these are monodominant communities of Anabasis salsa and communities of<br />

Atriplex cana.<br />

Table 1: Ecological-phytocoenotical dominant-determinant classification of European deserts.<br />

Subzone<br />

on the European<br />

Vegetation Map<br />

Northern lowland<br />

plain deserts<br />

Southern foothill<br />

deserts<br />

Formation Ecotype Association, complex of<br />

communities<br />

Artemisia lerchiana pl 1<br />

Caspian<br />

Lowland<br />

Artemisia lerchiana +<br />

pl hal Artemisia lerchiana-nabasis<br />

aphylla<br />

+<br />

hps Artemisia lerchiana-Poa bulbosa +<br />

hps Artemisia lerchiana-Stipa<br />

sareptana<br />

+<br />

hps Artemisia lerchiana-Stipa<br />

lessingiana<br />

+<br />

ps Artemisia lerchiana with<br />

Calligonum aphyllum<br />

+<br />

ps hal Artemisia lerchiana with<br />

Calligonum aphyllum and<br />

Krascheninnikovia ceratoides<br />

+<br />

Artemisia pauciflora pl hal Artemisia pauciflora +<br />

pl hal Artemisia pauciflora-<br />

Camphorosma monspeliaca<br />

+<br />

Artemisia arenaria ps Artemisia arenaria-Calligonum<br />

aphyllum<br />

+<br />

Anabasis salsa pl hal Anabasis salsa +<br />

Atriplix cana pl hal Atriplex cana +<br />

Artemisia fragrans pl Artemisia fragrans-Poa bulbosa,<br />

Catabrosella humilis<br />

Artemisia szowitsiana pl hal Artemisia szowitsiana-annual<br />

saltworts<br />

Salsola nodulosa pl hal Salsola nodulosa-Poa bulbosa,<br />

Catabrosella humilis with<br />

Artemisia fragrans<br />

Salsola ericoides pl hal Salsola ericoides-Poa bulbosa,<br />

Catabrosella humilis with<br />

Artemisia fragrans<br />

Salsola dendroides pl hal Salsola dendroides-Poa bulbosa,<br />

Catabrosella humilis with<br />

Artemisia fragrans<br />

Eastern<br />

Transcaucasus<br />

1 pl − pelitophytic, pl hal − pelitophytic halophytic, hps − hemipsammophytic, hps hal − hemipsammophytic<br />

halophytic, ps − psammophytic, ps hal − psammophytic halophytic<br />

Solonchak depressions are rather widely distributed in the Caspian Lowland. Aro<strong>und</strong> sors (solonchaks<br />

lacking vegetation) vegetation cover is formed by communities of hyperhalophytic annual saltworts<br />

and perennial saltworts, belonging mostly to the following formations: Halocnemum strobilaceum,<br />

Kalidium foliatum, Atriplex verrucifera, A. cana (Table 2).<br />

154<br />

+<br />

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